Tuesday, January 28, 2020

Making a Central Bank Independent

Making a Central Bank Independent Why has the idea of enhancing the independence of a central bank gained such popularity in recent years around the globe? Do accompanying accountability arrangements matter? The desirability of Central Bank Independence (CBI) has snowballed since (Alesina, A 1988) stated that his paper argues tentatively that independent Central Banks have been associated with a lower average inflation rate and may have been responsible for reducing politically induced volatility of monetary policy and inflation. As a result, we may be lured into the assumption that CBI was the brainchild of Alesina or Rogoff (who produced literature with similar results around the same time) and that it is a brand-new, groundbreaking concept. However, the issue of CBI is as old as central banking itself with David Ricardo arguing its benefits (or certainly the drawbacks of non-independence) in a paper written in 1824. Keynes articulated his thoughts on central bank independence while testifying to the 1913 Royal Commission into an Indian central bank. He stressed that the ideal central bank would combine ultimate government responsibility with a high degree of day to-day independence fo r the authorities of the bank. Clearly, as it is government legislation that created and gave powers to the central banks, there has always been a relationship between the two and they cannot be entirely distinct. Debate surrounding CBI considers the appropriate level of distinction (if any) and the potential benefits to the economy at large that such a separation would provoke. So if the theory behind the benefits of independence is almost two centuries old, then why has its popularity only soared in the last few decades? (Goodhart, C.A.E 1994) utilises Friedmans analysis of the Phillips Curve (1950s) to suggest that stagflation in the 1970s is a primary factor behind the surge towards CBI. The Phillips Curve displays the apparent inverse relationship (when the pressure of demand in an economy is low) between inflation and unemployment. Thus, Phillips suggested that the authorities were able to choose an optimal combination, or find a sufficient trade-off between the two, which is exactly what Governments attempted to do in the 50s and 60s. However, this theory was shot to pieces in the 1970s when the rate of inflation consistent with a given level of unemployment kept rising: stagflation. Friedman explained this by stating that the inverse relationship only ran true in the short-run. In the medium and long-run, he argued that the Phillips Curve would in fact be vertical and that there was no trade-off between inflation and unemployment. The implication of this was that those in charge could now use m onetary policy as an instrument to control inflation in the medium and long term without compromising growth or employment within the same time horizon; thus enabling fiscal policy and supply side measures to be utilised in stabilising shocks in the short term. Governments soon adopted medium-term financial strategies for bringing down inflation and began to use supply-side measures for promoting growth. Herein lay the problem, in that now a conflict of interest arose for those in charge. In the short-run, with expectations given, expansionary monetary and fiscal policies would raise employment and allow excess growth above the long-term level. However, Ministers were aware that although it may take some time to show up, higher inflation will be the inevitable result in the long-run. (Goodhart, C.A.E 1994) holds a very cynical view of politicians and suggests that they may forgo their medium-term economic responsibilities and lower taxes or raise expenditures before elections to ind uce a feel-good factor that would get them re-elected. The resulting inflation would only rear its ugly head after the election when they could tackle it by raising interest rates and thus maintain the boom/bust cycle. If they did not get re-elected it would be the next Governments problem and hence political short-mindedness and lack of credibility is laid bare for all to see. (Fraser, B.W 1994) is a lot less sceptical of politicians and argues that it is uncertainty rather than exploitation of the short-term inflation/unemployment trade-off that can inject inflationary bias into the policy making process. He argues that because no-one knows with any confidence what the long-term growth capacity limits are or what the natural rate of unemployment is, it is very difficult for politicians to heed warnings about operating above these limits whilst under pressure from the electorate to maintain or stimulate growth. Similarly, they do not know the length of the lags between policy chang es and their impact on growth and inflation, thus Fraser implies that it is only natural for politicians to believe that they can push the economy a little bit further. Or, as William McChesney-Martin, the Governor of the US Fed from 1951 to 1970, said They may be reluctant to take away the punchbowl just when the party gets going. Irrelevant of your personal degree of cynicism towards politicians, it is clear that an independent authority with a long-term vision of price stability and no inclination towards inflation is the only remedy to rescue the electorate from a spiral of inflationary doom inflicted by governments. Pre 1971 a large degree of price stability was autonomous in the developed world with first the Gold Standard, then the Bretton-Woods system anchoring prices to a fixed level. (Fraser, B.W 1994) proposed that the earlier arrangements had imposed an international discipline on countries but when those arrangements passed into history, the responsibility for maintaining price stability reverted to national authorities. This perhaps adds more credibility to Frasers lack of scepticism as for example, (excluding war times) the UK government had not had this burden since 1717, a completely incomparable financial era. Coupling this 250 year knowledge gap, with the temptation of short-term benefits at long-term costs, it is of no surprise that Government-managed monetary policy was doomed to collapse. After this was realised, the move towards an Independent Central Bank evolved naturally as it solved both the politicians and publics concerns. Following over a decade of failure, politicians wanted rid of the price stability burden and someone else to blame for its failure (Kane, E 1980) and the public wanted monetary policy to be controlled by an institution with credibility (Rogoff, K 1985) so that their expectations were met. Despite conveniently tying together, these two arguments are not exhaustive in explaining the rise in popularity of CBI in recent times. Under the Maastricht Treaty, all states wishing to enter the European Union must have an independent national central bank so as to complement the ECB and the European System of Central Banks, whose job it is to ensure that the Euro area benefits from price stability. The EU has adopted such a hard-line stance on the basis that the success of the German economy of maintaining low inflation has arisen from the independent nature of the Bundesbank. Therefore, the installation of an independent central bank (the scope of independence is not legislated) has been forced upon all twenty-seven member states regardless of whether they had previously suffered inflationary problems. However, it must be noted that by applying to join the EU, each member state is already willing to concede its control over monetary policy in the long-term to the ECB, so an inte rmediate step to a national independent central bank would not present a significant hurdle. Many commentators have looked at who in the economy benefits most from CBI and have drawn conclusions on the reason for the growth of central bank independence from there. Those emphasising the interests of the financial sector as key are perhaps the most logical. (Posen, A 1993) and (Bowles, P White, G 1994) suggested that independence has been encouraged by financial interests and global institutions taking advantage of a crisis of governance in the 1980s and 1990s. The benefits for such institutions are obvious; a credible monetary policy allows for accurate expectations within a business plan and low inflation maintains real wages and ensures low interest rates for accessing credit. Therefore we can accredit some popularity of central bank independence to the shift in political power towards large corporations, with the most notable example being the USA. Linked to this, (Maxfield, S 1997) proposed that Governments in some fast-growing economies hold the perception that foreign investment from such large corporations will therefore be more forthcoming if they have an independent central bank. Other commentators have looked elsewhere in society with (Piga, G 2000), suggesting that the aging of some populations has promoted creditor interests. However, this is not supported by the speed of reform as although populations are aging, they do not do so suddenly whereas the popularity of central bank independence took off very rapidly. So it can be seen that there may be many different reasons behind the surge in popularity of CBI and as a result it seems natural to conclude that different levels or types of independence would be more suitable for these different variations. Similarly, the different nature of governance and democracy in countries dictates the need for a reasonable amount of flexibility and varying accountability within central bank independence. The norm within the literature is to follow Fischers (1994) dichotomy between goal and instrument independence, although many different measures of independence have been investigated and published; most notably (Cukierman, Webb, and Neyapti, 1992) and (Grilli, Masciandaro, and Tabellini, 1991). Goal independence refers to the central banks capacity to choose policy goals without being under the direct influence of the fiscal authority (usually the Government). The Bank of England lacks goal independence because the inflation target, which is very specific measure, is set by the government. In the USA, the Humphrey-Hawkins Act requires the Federal Reserve to conduct monetary policy to promote the goals of maximum employment, stable prices, and moderate long-term interest rates. These goals are described in vague terms providing the Fed some leverage to translate these into operational goals and thus allowing it a high level of goal independence. Instrument independence alludes to the central banks ability to freely adjust its policy tools in pursuit of the goals of monetary policy (Walsh, C 2005). Despite lacking goal independence, the Bank of England has instrument independence: it is provided its inflation mandate by the government and then it is able to choose its instruments without any further direction. However, the Federal Reserve has complete instrument independence in addition to having a large degree of goal independence. How can a nation that prides itself for being democratic justify handing over complete control of monetary policy to a group of un-elected officials? In addition, such a system would surely not resolve the issue of uncertainty regarding inflation policy as the public, who clearly distrusted them before, now have to rely on politicians to choose suitable people to control monetary policy. The simple solution is accountability; if central banks make their decisions transparently and/or are held acco untable for their actions, the public can feel a lot more confident in making expectations. The reputation of a central bank plays a key role in how much accountability is required; the longer the CB has delivered its promises/targets, the more trustworthy it is deemed to be and the less accountability is required. The empirical evidence seems to support this assertion. The German Bundesbank, often compared to Rogoffs Conservative Cental Banker (Rogoff, K 1985), has a very high level of independence but almost no accountability and this is sustainable only because of its reputation. Since the hyperinflation of the 1920s the Bundesbank has been careful to reflect, or even cultivate a public acceptance of the need for price stability (Bank of England 1996). This means that the inflation-averse German people are happy to trust the Bundesbank to deliver low inflation because of its impeccable track record over the last 50 years. This suggestion is strengthened by the following graph, where low inflation is related positively to low accountability: At the other end of the spectrum, the Bank of England and even more so the Reserve Bank of New Zealand are held accountable for their results, despite their comparative lack of independence. This may be explained by their relative infancy within the realms of CBI (The BoE became independent in 1997 and the RBNZ in 1989) and so in either case there has not been enough time to build a reputation. With regards to the UK, the terrible collapse of the Medium-Term Financial Planning system under Thatcher may remain a coal stoking the fire of the publics political cynicism, adding further need for a high level of accountability. There are three main channels by which the BoE demonstrates its transparency and accountability to both the Government and the public at large. Primarily, the inflation target itself is the cornerstone of the authorities medium-term price stability objectives and provides an indisputable measure of failure or success that is simple to understand. This is in stark contrast to the MTFS where numerous measures (e.g. Â £M3, M1, PSL2), which were not immediately recognisable to the man on the street, were used and muddied the water if targets were missed. In addition the minutes of the meetings between the Chancellor and Governor, where monetary policy decisions are made and discussed, are published each month along with the Inflation Report, detailing the Banks own scrutiny of inflationary patterns. In comparison to The New Zealand Approach, where the Governor can be sacked for missing an inflation target, having the Governor of the BoE write a letter for the same crime seems particular ly soft. However, if we compare the relative successes of the two banks since they adopted independence, we see that until 2007 the BoE never missed a target and that in 1990 New Zealand had 8% (RBNZ Website) inflation when its target was 0-2%. This evidence seems to add credence to the suggestion that more accountability is required with a lesser reputation. The 1990s saw both developed and developing countries move in their droves towards increased central bank independence. This trend was sturdily influenced by empirical analysis of the relationship between macroeconomic performance and independence [see Alesina and Summers (1993), Jonsson (1995), andEijffingeret al. (1998)], which among the developed countries suggested a negative relationship between independence and inflation. For this reason alone it is of no surprise that CBI popularity grew, but coupled with the earlier stated reasons it seems to have become a necessity to successfully run a modern Government and economy. By adopting independence, a restriction on government interference in monetary policy is imposed; while making the central bank transparent and accountable imposes a restraint on how it utilises this independence. Both of these constraints are desirable as they allow those more knowledgeable to influence policy and provide those responsible for making policy someone else to blame if it fails. However, transparency by itself is not necessarily adequate for a monetary institution; after all, what good is the CB missing its objectives but just being very honest about it after? Instead, transparency can help the institution combat inflation bias and promote confidence in expectations, either by itself or in conjunction with central bank independence or even a formal central bank contract (Bank of England 1996), as in New Zealand. Finally, the empirical evidence linking independence, accountability and low inflation is conclusive. From the graphs above we note the inverse relationship between accountability and independence, which suggests that they are substitutes rather than compliments (Bank of England 1996). Hence, we can conclude that in terms of inflation targeting, accountability is equally as important as central bank independence, as one or the other (not necessarily both) is required for success. In addition the positive relationship between accountability and inflation history suggests that, at least for a short time period, accountability can be used as a substitute for a reputation of low inflation. Thus providing an instantaneous removal of the problems associated with the rational expectations model and allowing low inflation to be enjoyed by all. References: Alesina, Alberto. Macroeconomics and Politics. In NBER Macroeconomics Annual, pp. 17-52. Cambridge, 1988. Alesina A, Summers L.Centralbankindependenceand macroeconomic performance: some comparativeevidence. In Journal of Money, Credit and Banking 25: pp 151-62(1993) Bank of England. Central Bank Independence Accountability: Theory and Evidence. In Bank of England Quarterly Bulletin, February 1996, pp-63-68 Bowles, P and White, G. Central bank independence: A political economy approach.In The Journal of Development Studies31(2) (1994), pp. 235-264 Cukierman, A., S. B. Webb, and B. Neyapti. Measuring the Independence of Central Banks and its Effects on Policy Outcomes. In The World Bank Economic Review, 6, pp. 353-398. 1992 Fischer, S. Modern central banking. In The future of Central Banking (1994), Eijffinger SCW, Schaling E, Hoeberichts M.Centralbankindependence: a sensitivity analysis. In European Journal of Political Economy 14: pp.73-88 (1998) Fraser, B.W. Central Bank Independence: What does it mean? In Reserve Bank of Australia Bulletin, 1994. Grilli, V, D. Masciandaro, and G. Tabellini. Political and Monetary Institutions and Public Financial Policies in the Industrial Countries. In Economic Policy 6, pp 341-392. 1991 Goodhart, C.A.E. Central Bank Independence. In Journal of International and Comaparative Economics, 3. 1994. Jonsson G. Institutions and macroeconomic outcomes theempiricalevidence. In Swedish Economic Policy Review 2: pp.181-212. (1995) Kane, E. Politics and Fed Policymaking: The More ThingsChange, the More They Remain the Same,In Journal of Monetary Economics,6,2(April 1980), pp. 199-211 Maxfield, S. Gatekeepers of Growth: The International Political Economy of Central Banking in Developing Countries 1997 Piga, G. Dependent and Accountable: Evidence from the Modern Theory of Central Banking,In Journal of Economic Surveys vol. 14(5), December 2000, pp 563-95 Posen, A. Why Central Bank Independence Does Not Cause Low Inflation: There is No Institutional Fix for Politics. In R. OBrien (ed.), Finance and the International Economy. 1993, pp 40-65. Rogoff, K. The optimal degree of commitment to an intermediate monetary target.In Quarterly Journal of Economics100(1985), pp. 1169-1189 Walsh, C. Central Bank Independence Prepared for The New Palgrave Dictionary December 2005 RBNZ Website: http://www.rbnz.govt.nz/keygraphs/Fig1.html

Monday, January 20, 2020

Changes in Latitude :: Personal Narrative Essays

Changes in Latitude I was nominated to go to Singapore based on my achievements in the science fair. However, looking back on my recent experience in Singapore, it was the people, and not the science, that really stick in my memory. I think that's just as well. Most of the science will be yesterday's news by the time I go to college anyway. This was my first opportunity for international travel, and certainly, Singapore is about as far away as it is possible to travel from Pennsylvania. Singapore is also far away from home in its culture and climate. Traveling there opened my eyes to new ideas, values, and attitudes. The talks at the festival were great. One of the highlights of the festival for me was the spirited and humorous talk given by the Australian scientist / radio show host. I also learned a lot of new information about the human genome project, which seemed to be a prime interest of APEC's scientific community, and certainly an important project for demonstrating cooperation. Several distinguished scientists talked to us about their work with HUGO and their role in deciphering the genetic code of the human species. In addition to this exposure to scientific research in the region, I learned something about the education system of Singapore. It is, I found, somewhat different than ours. They swap the last two years of high school for something called junior college. These students are looked on with admiration by the younger students, and the adults from Singapore seem to entrust all leadership responsibilities to them, rather than the younger students. These junior college students from Singapore were also favored over students of the same age from other countries. This was upsetting to some. For example, I know one junior college-aged boy in my group who was not from Singapore complained of this. I understand, however, that this preference for the junior college students from Singapore reflects their culture's reverence for elders. Besides learning something about the education in Singapore, I had the opportunity to discuss international careers with other attendees of APEC. I think that the biggest advantage of thinking "internationally" is that you benefit from having minds from all over the globe help you solve scientific problems. And from a business perspective, you can make use of a global market to sell to a wider variety of customers.

Saturday, January 11, 2020

Chinese Cinderella Response to Lit

Big Brother and Second Brother Two of Deadline's brothers are malicious and mischievous. Why you say? Well, because clues in the context of Chinese Cinderella tell me so. I can tell you that Deadline's eldest brother is mischievous from Just two quotes of the book. One Is, â€Å"Silently but swiftly, Big Brother suddenly approached Ye Ye and carefully pinched the nasal hair between his forefinger and thumb. † This Illustrates that Big Brother thinks It's funny to pull out one of his grandfather's nose hairs while his senior Is leaping.Another quote Is, â€Å"Laughing hysterically, Big Brother rushed out of the room, slid down the banister and made a clean getaway Into the garden, all the time holding Ye Yes's hair aloft Like a trophy. † This quote shows that Big Brother Is so mischievous that he would even pluck out a nasal hair of his grandpa's and even evade his old man's anger for the fun of it! Second Brother also has a very distinct personality. He is malicious to Deadline because of jealousy and everything that goes wrong for him.To support my claim, ere are two quotes and why I chose these quotes. â€Å"Deliberately he took my right arm under the table and gave it a quick, hard twist while no one was looking. † Second Brother is so cruel that he hurts Deadline right at the dinner table where anyone may see him. â€Å"n. You ugly little squirt! This'll teach you to show off your medal! † Deadline's brother openly shows that he acts maliciously not Just for no reason, but also because he is Jealous of the attention that Deadline is getting because of an award she received.

Friday, January 3, 2020

Pastoral Landscapes - Free Essay Example

Sample details Pages: 21 Words: 6264 Downloads: 2 Date added: 2017/06/26 Category Management Essay Type Research paper Did you like this example? Hedgerow Management in Pastoral Landscapes Abstract Hedgerows are an important part of the British landscape, providing both food and shelter for a number of taxa. As part of the UK government’s Environmental Stewardship (ES) Scheme, farmers are granted subsidies for, amongst other things, ‘enhanced hedgerow management’. Although hedgerow management under ES is expected to have beneficial effects for taxa such as invertebrates and birds, less is known about the effects ES management will have on small mammal communities. Don’t waste time! Our writers will create an original "Pastoral Landscapes" essay for you Create order The aim of this study was to investigate whether this ‘enhanced hedgerow management’ is affecting hedgerow characteristics in pastoral landscapes and whether small mammal abundances are increased under ES managed hedgerows. ‘Conservation buffer strips’ (2m+ unimproved grassy margins) were investigated as a possible improvement to ES hedgerow management. Using live trapping methods, I investigated small mammal abundances in ES managed hedgerows compared with non-ES managed hedgerows. Wood mice Apodemus sylvaticus and bank voles Clethrionomys glareolus were the most abundant species, with some captures of field voles Microtus agrestis and common shrews Sorex araneus. Small mammal abundances were increased in ES managed hedgerows, however, the presence of a ‘conservation buffer strip’ was more significant in increasing small mammal densities. ES management showed no definite effect on the hedgerows’ characteristics. Introduction Agricultural intensification since the 1940s has led to widespread and significant reductions in the biodiversity of many agricultural areas. This drive for greater yields has been linked with the population decreases seen in many species of farmland specialists and non-specialists who often inhabit farmland (Robinson and Sutherland, 2002). Farmland habitats can be categorised into non-linear habitats such as set-aside, cropped fields and woodland areas, and linear habitats, generally field boundaries, such as ditches, banks, streams and hedgerows. These field boundaries remain relatively undisturbed areas and are therefore significant wildlife corridors within otherwise inhospitable agricultural landscapes (Tew, 1994). Although there continued to be a reduction in total area of hedgerows within the UK during the 80s and early 90s, the last decade has seen small increases in the area of hedgerow as their conservation significance became more documented (Barr and Gillespie, 2000) . This increase in the number of hedgerows has been driven by government backed grants. Countryside Steward (CS), set up in 1991 encouraged selected farmers to enhance and conserve the wildlife within their farms, a large part of this involved the laying of new hedgerows. The CS schemes have now been superseded by the Environmental Stewardship Schemes. More recently, hedgerow grant pilot schemes have been set up in a number of regions to encourage landowners, both farmers and non-farmers to manage their hedgerows more effectively; these grants are available to pay for gapping up, hedge laying or coppicing. Small mammals in pastoral land are largely confined to hedgerows or other non-crop features and are therefore particularly vulnerable to intensification (Bates and Harris, 2009). Small mammal species constitute the main prey biomass for a number of species of mammals and birds, and therefore small mammal abundance directly influences the abundance and diversity of predator spec ies contributing to the complexity of local food webs (Korpimaki and Norrdahl, 1991). There remains some debate on the importance of linear habitats for small mammals, with some suggesting that they cannot support viable populations, that those found in hedgerows are ‘sink’ populations (Tattersall et al. 2004). However, there is evidence that small mammal abundance and diversity does not depend on the linear or non-linear character of the habitat and that linear habitats can support viable populations (Gelling et al. 2007). Thus, in large expanses of uninhabitable grassland, field boundary hedgerows are of great importance for maintaining small mammal populations in an agricultural landscape, but differing farming practices can lead to a huge variety in the quality of these habitats As the emphasis of farming has shifted there have been a number of agri-environment schemes introduced across Europe with the aim of reversing the effects of previous intensification a nd enhancing agricultural land for wildlife (Kleijn and Sutherland, 2003). The UK introduced a new set of farming standards in 2005 with farmers now guaranteed subsidy payments, known as ‘cross-compliance’, as long as they follow a set of prescribed conditions aimed at improving the environmental value of their farms. A compulsory code of good practice will preclude farming land within 2 m of the centre of a hedge (DEFRA, 2005a). Beyond cross-compliance subsidies, farmers can also apply to put their farmland into Environmental Stewardship (ES). ES is a tiered system, with Entry-Level ES designed to allow most farmers access to the payments by compiling a farm management plan that aims to improve the features of their farm for wildlife and to maintain/improve the scenic value of the British countryside. The enhanced hedgerow management option within ES requires that the farmer cut the hedge no more than once every 2 years, that hedgerows are cut during the winter and tha t cutting be staggered across the farm. The combined aim of these prescriptions is to ensure that at least some of the hedgerow is allowed to flower every summer (Defra, 2005b). Properly managed hedgerows are valuable features, playing a key role in enhancing the wildlife value of farmland. Flowering hedgerows are an important source of food and shelter for a number of birds (Hinsley and Bellamy, 2000). Studies suggest that the ES schemes will have a beneficial effect, mainly for taxa such as invertebrates and birds (Kleijn and Sutherland 2003), Whittingham (2007) emphasizes the importance of monitoring the effects of ES to ensure that the scheme’s prescriptions meet the needs of a greater range of species. It is much less well understood how the changes to hedgerow management will effect small mammal abundance, and it is important that there is greater understanding of the factors that influence small mammal populations since small mammals provide the major source of prey biomass for many larger predators (Love et al., 2000). Small mammals also play a role in a range of important ecosystem processes (Hayward and Phillipson, 1979). Previous studies have established the main effects of varying hedgerow management within arable landscapes (Shore et al. 2005). Arable environments provide cover for small mammals due to the height and density of the crop. Small mammals have been shown to make substantial use of the field at certain times of the year (Tattersall et al. 2001; Tew et al. 2000; Todd et al. 2000). However, no small mammal species have been shown to make use of agriculturally improved pastoral fields at any time of year (Montgomery and Dowie 1993). Grazed pastoral land provides very little cover, restricting the movements of resident small mammal communities. Therefore, hedgerow management in predominantly dairy and cattle areas will likely have a large influence on the success of small mammal populations (Gelling et al. 2007). In particular , the level of ground cover vegetation along the hedgerow and the presence of some form of non-farmed margin can significantly affect the small mammal abundance (Bates and Harris 2009, Gelling et al. 2007). The 2m margin prescribed by ‘cross compliance’ is irrelevant in terms of providing cover within pastoral landscapes. Although the 2m margin remains uncut and clear of interference from the farmer (no fertilisers), year round grazing will mean that little cover is offered right up to the base of the hedgerow. Therefore, whereas ES management may boost small mammal numbers within arable areas (Shore et al. 2005), the value of ES hedgerow management within pastoral landscapes is less well understood. I utilised a number of hedgerow sites to compare hedgerow structure and small mammal communities on ES farms versus non-ES farms. For each farm, one site was selected to be representative and one to include a significant (2m plus) conservation buffer strip of unimproved, no n-grazed grass/shrubland. I aimed to investigate (i) how ES management effects the hedgerow characteristics, in particular the level of ground cover for small mammals (ii) whether these ES prescriptions are providing any significant benefit for small mammal densities and (iii) as the movements of small mammals within pastoral landscapes are so restricted, could small mammal assemblages in hedgerows be significantly improved by including an unimproved, non-grazed, grassy margin or ‘conservation buffer strip’ (2+m from the edge of the hedgerow). Methods Sites The study was conducted over 20 different farms spread across County Durham and Northumberland. The farms were selected due to their suitability for this study, each farm containing both a hedgerow site with a conservation buffer strip and at least one without. All farms selected were representative in terms of habitat of those within the local area. A hedgerow was defined as a continuous line of woody vegetation no more than 3m tall. Hedgerow Survey The farms were paired, with one ES farm neighbouring a non-ES farm, making 10 farm pairs and 20 farms in total. Hedgerow surveys were carried out throughout June 2009. 10 hedgerows were randomly selected on each farm. All hedgerows on each farm were surveyed using an edited version of the Defra Hedgerow Survey Form and handbook (DEFRA, 2007). Each hedgerow was measured to determine its cross-sectional area. The character of the hedgerow was scored by reference to a series of standard diagrams, noting the level of available ground level cover for small mammals (1=little or no vegetation cover at ground level, 2=gappy cover at ground level, 3=constant vegetation cover from hedgerows at ground level). Additional variables were recorded, including whether the hedge had been flailed (mechanically cut) recently, i.e. during the previous winter, the number of standard and veteran trees and the number of woody species within the hedgerow. The data sets for cross-sectional area, level of gr ound vegetation cover and the number of woody species were averaged to produce an overall mean value for each farm. The number of flailed hedgerows was summed to give an overall percentage of hedgerows flailed on each farm. Trapping Procedure Previous trapping studies have shown that, unlike in arable land, small mammals within pastoral land stay almost entirely within the hedgerows and therefore hedgerows can be treated as linear habitats (Gelling et al. 2007). Trapping was carried out in two major trapping sessions, mid-April to June and mid-July to August, 2009. Within each of the 20 farm sites I selected a representative hedgerow and a hedgerow flanked by an unimproved 2m+ grassy margin, designated a conservation buffer strip, making a total of 40 trapping sites. Where possible the hedgerow sites were selected randomly, however, each ES site was required to have been managed according to the prescriptions of Stewardship farming, i.e. the hedgerows were cut not more than once every two years and the farmers adhered to the prescribed 2m margin of non-interference (2m from the centre of the hedge) (DEFRA 2005a, DEFRA 2005b). Every hedgerow selected was flanked by improved or semi-improved grassland for the grazing of d airy cattle and/or the production of silage. At each site, a 104m section of isolated hedgerow (not directly connected to woodland) was selected.13 Longworth traps were placed at ground level within the hedgerow, at 8m intervals. Traps were provisioned with hay, apple, oat grains and dried mealworm. The traps were set at dusk and checked at dawn and dusk for three days. All targeted animals that were captured were fur-clipped to help identify re-captures. Species, sex and weight were recorded for each animal before release at the point of capture. Analysis Hedgerow characteristics were recorded and analysed using a paired measures multivariate analysis of variance (MANOVA) (SPSS 17.0.2). I had multiple dependent variables that I wished to analyse, however, using multiple one-way ANOVAs to try to do this would have raised the probability of a Type I error (Gibson et al. 2007). Therefore the data was investigated using a MANOVA which controls the experiment-wide error rate. Multiple dependent variables that were related (e.g. Cross sectional area of hedge and amount of ground cover, etc.) were analysed in one test, with the hedgerow management (ES managed or non-ES managed) being treated as the two levels of the treatment factor (Gibson et al. 2007). There was a total of 4 dependent variables; the mean cross-sectional area, the percentage of flailed hedgerows, the average number of woody species and the mean level of ground cover. For each trapping session the relative density was estimated as the minimum number alive (MNA), or the total number of individuals caught over the three days. Species richness was calculated as the number of different species caught. Using General Linear Modelling (GLM; Minitab 15), I examined the relationships between small mammal densities and a number of predictor variables. The dependent variables I investigated were the overall total small mammal density (MNA) and the total biomass of all small mammals caught within 104m. I also investigated the density of each individual species, constructing similar models for the number of captures and biomass for each individual species. I focused on wood mice Apodemus sylvaticus and bank voles Clethrionomys glareolus. There were also some captures of field voles Microtus agrestis and common shrew Sorex araneus, these data were not investigated individually but were included in the total density of small mammals and the total biomass. The predictor variables considered were the presence/absence of ES management, the presence/absence of a con servation buffer strip and the number of standard and veteran trees within the hedgerow. The relationships were analysed using a backward stepwise GLM, with all main predictors and their first order interactions initially included within the model. The insignificant interactions were then removed. Each trapping session was carried out over 3 days on 4 sites on neighbouring farms, the variation between trapping locations and times was taken into account by including the variable ‘block’ within the initial model, however, it was found to have no significance and was therefore removed from the final model. There are well documented seasonal variations in small mammal abundance (Alibhai and Gipps 1985; Flowerdew 1985; Butet et al. 2006), therefore, as there were two major trapping seasons (mid-April to May and Mid-June to July) I included the variable ‘season’ in all models. The number of captures of field voles and common shrew were too low to allow thorough a nalysis; however, the number of captures for each species was investigated using a Kruskal-Wallis test (Minitab 15) to determine the relationship between the presence of a buffer strip and their individual abundance. Results The total number of catches was 276 individual small mammals of four different species, during 240 trap sessions (dusk till dawn and dawn till dusk). The most abundant species were wood mice, making up 45% of the captures, 11% of which were juveniles, with a total capture of 122 individuals (61 in the first season of trapping and 61 in the second season). 32% (89 individuals) of captures were bank voles, none of which were juveniles, with 26 captures in season 1 and 53 captures in season 2. 17% of captures (48 individuals) were common shrews and 6% (17 individuals) were field voles. Table 1. Summary of the number of captures for each species Total Wood mice Captured – Season 1 (juveniles) / Season 2 (juveniles) Bank vole – Season 1 / Season 2 Field vole – Season 1 / Season 2 Common shrew – Season 1 / Season 2 Total – Season 1 / Season 2 Total N trapped throughout study 122 – 61 (2) / 61 (11) 89 – 36 / 53 17 – 4 / 13 48 – 28 / 20 276 – 129 / 147 Percentage of total 44 33 6 17 100 Percentage of hedgerows present 93 46 23 45 Effect of ES Management and Buffer strips A total of 40 hedgerows were surveyed with 20 hedgerows under ES hedgerow management and 20 hedgerows under non-ES management. ES sites had been under ES hedgerow management for 2 years or more. The measured dimensions of the hedgerow were used to estimate the hedgerow cross sectional area. Analysis using a paired measures MANOVA found no significant difference in the size of ES managed hedgerows to the size of non-ES managed hedgerows (F(1,9)=0.847, P=0.381). ES management also had no significant effect on the percentage of flailed hedgerows within the farm (F(1,9)=0.019, P=0.889). The woody species diversity within hedgerows was not significantly different between ES managed hedgerows and non-ES managed hedgerows (F(1,9)=3.047, P=0.115). There was a significant positive association of the presence of ES hedgerow management with the level of woody vegetation cover at ground level (F(1,9)=10.613, P=0.010). Table 2. Comparisons of hedgerow characteristics on ES managed farms versus non-ES managed farms. Data were analysed using a paired MANOVA. Mean (SE) Hedgerow characteristic Description of measurement ES Non-ES F(1,9) P Area Average cross sectional area/m2 2.99 (0.12) 2.83 (0.14) 0.847 0.381 Flailed Percentage of hedgerows that had been recently flailed (flailed during previous Winter) 26.00 (2.21) 25.00 (6.54) 0.019 0.893 Species diversity Number of woody species 3.16 (0.24) 2.73 (0.27) 3.047 0.115 Small mammal cover Average Area of small Mammal cover (1=little or no vegetation cover at ground level, 2=gappy cover at ground level, 3=constant vegetation cover from hedgerows at ground level) 2.63 (0.87) 2.13 (0.11) 10.613 0.010 Small Mammal Assemblages Backward stepwise general linear modelling was used to analyse the data. The results showed that buffer strips have a significant effect on the total number caught within the hedgerow (F(1,35)= 16.29, P0.001), with the numbers of captures rising along hedgerows flanked by buffer strips. ES management also appeared to have a positive significant effect on the total number of captures (F(1,35) = 5.23, P=0.028), however, the positive association with increased captures was not as strong as seen with buffer strips. The number of standard trees did not significantly effect the total number of captures (F(1,35)=0.91, P=0.346). Season had no significant effect on the number of catches (F(1,35) = 1.09, P=0.305), and there were no significant interactions between variables affecting the number of captures. A GLM for total biomass showed similar results with Season (F(1,34)=0.83, P=0.369) and the number of standard trees (F(1,34)=1.12, P=0.298) both having no significant effect on the tot al biomass. ES management had a positive association with total biomass (F(1,34)=4.92, P=0.033), as did the presence of a buffer strip (F(1,34)=27.62, P0.001). Interestingly, there was an interaction between Season and the presence/absence of a Buffer Strip which appears to have a significant effect on the total biomass (F(1,34)=3.18), P=0.083), with greater total biomass found within hedgerows flanked by buffer strips in the second season of trapping (mid July-August). Wood mice were the most common species trapped, contributing 45% of the captures. The factors affecting wood mice captures were analysed using a backward stepwise GLM. Season had no significant effect (F(1,34)=2.36, P=0.134). Unlike the model involving ‘total captures’, ES management (F(1,34)=0.07, P=0.798) and Buffer Strip (F(1,34)0.01, P=0.947) had no significant effect on the number of wood mice captured. The results show that number of trees within a hedgerow is the most significant factor affecti ng wood mouse abundance (F(1,34)=79.65, P0.001). There was also an interaction between the season and the number of trees within a hedgerow which had a significant effect on the number of wood mouse captures (F(1,34)=4.81, P=0.035). The number of wood mouse captures was significantly increased in hedgerows containing a greater number of trees in the second season of trapping (mid-July to August). A backward stepwise GLM constructed for total wood mice mass showed similar results. Season had no significant effect (F(1,35)=1.36, P=0.252). ES management had no significant effect on the total wood mice mass (F(1,35)=0.26, P=0.616). The presence/absence of a buffer strip also had no significant effect on the total mass of wood mice (F(1,35)=0.05, P=0.831). However, the number of trees within a hedgerow was shown to have a strong positive association with the total mass of wood mice (F(1,35)=49.03, P=0.003). A backward stepwise GLM was constructed for both ‘bank vole capturesâ₠¬â„¢ and ‘the total bank vole mass’, both models produced similar results. Season had no effect on bank vole captures (F(1,35)=2.06, P=0.160) and total bank vole mass (F(1,35)=1.66, P=0.206). The presence of ES management on the hedgerow had a significant positive effect on the number of bank vole captures (F(1,35)=7.15, P=0.011) and on the total bank vole mass (F(1,35)=5.91, P=0.020). The presence of a buffer also had a significant effect, increasing the number of bank vole captures (F(1,35)=34.90, P0.001) and the total bank vole mass (F(1,35)=28.11, P0.001). The number of standard and veteran trees also appeared to have significant effect on bank vole captures (F(1,35)=4.41, P=0.043), bank vole abundance is reduced in areas with more veteran trees. However, the total bank vole mass was not significantly effected by the number of veteran trees (F(1,35)=2.32, P=0.137). Table 3. Summary statistics from general linear models Model Variables F P Adj. R2 Total Captures Season F(1,35)=1.09 0.305 53.79% ES Managed F(1,35)=5.23 0.028a Buffer Strip F(1,35)=16.29 0.001a Standard Trees F(1,35)=0.91 0.346 Total Biomassc Season F(1,34)=0.83 0.369 65.32% ES Managed F(1,34)=4.92 0.033a Buffer Strip F(1,34)=27.62 0.001a Standard Trees F(1,34)=1.12 0.298 Season*Buffer Strip F(1,34)=3.18 0.083b Wood Mice Captures Season F(1,34)=2.36 0.134 79.72% ES Managed F(1,34)=0.07 0.798 Buffer Strip F(1,34)0.00 0.947 Standard Trees F(1,34)=79.65 0.001a Season*Standard Trees F(1,34)=4.81 0.035a Total Wood Mice Massd Season F(1,35)=1.36 0.252 69.06% ES Managed F(1,35)=0.26 0.616 Buffer Strip F(1,35)=0.05 0.831 Standard Trees F(1,35)=49.03 0.003a Bank Vole Captures Season F(1,35)=2.06 0.160 54.76% ES Managed F(1,35)=7.15 0.011a Buffer Strip F(1,35)=34.90 0.001a Standard Trees F(1,35)=4.41 0.043a Total Bank Vole Masse Season F(1,35)=1.66 0.206 50.74% ES Managed F(1,35)=5.91 0.020a Buffer Strip F(1,35)=28.11 0.001a Standard Trees F(1,35)=2.32 0.137 a – Significant to the 95% confidence level b – Significant to the 90% confidence level c Total Biomass was square root transformed before analysis. d Wood Mice Mass was square root transformed before analysis. e Bank Vole Mass was square root transformed before analysis. A total of 17 field voles were captured, with all 17 trapped in hedgerows flanked by a conservation buffer strip. A total of 48 Common shrews were trapped, 81% of which were caught in hedgerows not flanked by a buffer strip Table 4. Non-target species captures. Effect of buffer strip, analysed using Kruskal-Wallis test. Total Captures Species Buffer Strip Present No Buffer Strip H P (adjusted for ties) Field vole 17 0 8.30 0.004 Common shrew 9 38 12.73 0.001 Discussion Hedgerow characteristics are known to affect small mammal numbers. Hedgerows with many gaps and a lack of ground cover support significantly lower small mammal populations (Gelling et al. 2007). Small mammals will select against hedgerows with a lack of vegetative cover due to the increased risk of predation (Orrock et al. 2004). Our results suggest that ES farms produce denser hedgerows with more cover at the ground level than non-ES farms. This is reflected in the small mammal survey which shows a somewhat strong association between small mammal numbers and ES hedgerows. However, having surveyed the farms and the farmers, I acknowledge that a wide number of variables affect the characteristics of the hedgerow. I suggest that the state of the hedgerows for small mammals is more significantly affected by the mindset of the farmer. Those farmers who have moved onto the Entry level ES scheme are generally those who most actively manage their farm. One supporting piece of data for thi s theory, is the number of flailed hedgerows on ES farms compared to non-ES farms. The hedgerow survey found that there were no differences in the number of recently flailed hedgerows within ES farms compared to non-ES farms, therefore, even though the cutting of hedgerows on ES farms is restricted, it still occurs as often on the ES farms within this survey than on the non-ES farms. The suggestion is that those farmers who are on the ES scheme are more actively involved in managing their farm, including their hedgerows, therefore hedgerows on ES farms commonly provide denser vegetation, less gaps and more cover at ground level. The typical ES farmer is more actively managing the hedge as a boundary or barrier to cattle than the typical non-ES farmer. The author suggests this conclusion having discussed hedgerow management with the farmers as part of the hedgerow survey and having a background in agriculture, however, it is also recognised that this topic goes beyond the scope and a vailable data of this investigation. Hedgerows can be thought of as corridors linking woodland habitat, allowing small mammal migration (Soule and Terbough 1999), however, within the British pastoral landscape, hedgerows are often acting as the sole habitat for small mammals (Fitzgibbon 1997). My investigation found that the ratio of juvenile to adult wood mice increased during the season, with greater numbers present later in the summer, this is consistent with the observations of others (Alibhai and Gipps 1991, Flowerdew 1991). The breeding season for most small mammals begins in spring and ends in late summer, therefore it is natural that more juveniles are present in hedgerows as the summer progresses and they travel outward to establish their own home ranges. The presence of fully grown, breeding adults in both seasons of trapping indicates that animals are resident within the hedgerows, providing support for the argument that linear habitats can provide suitable habitat to support viable populations of small mammals. My results show that the total small mammal abundance and therefore the availability of prey biomass for predators is increased in hedgerows under ES management. The results of the hedgerow survey suggest that there is greater ground level vegetation cover in ES hedgerows. An increase in the amount of physical habitat creates greater foraging opportunities and can increase small mammal abundance (Gelling et al. 2007). Small mammals prefer hedgerows with greater ground level cover as they provide better refuge from predators (Orrock et al. 2004). Whereas the benefits of ES management for small mammal abundance remain unclear, this investigation highlights the importance of buffer strips. The value of unimproved grassy margins, in arable landscapes, for small mammal numbers has already been shown (Shore et al. 2005). This study suggests that the presence of a buffer strip along a hedgerow can provide a much improved habitat to support larger small mammal numbers in hedgerows within pastoral landscapes. Grassy margins are a refuge for small mammals beyond the hedgerow; they allow increased safety for foraging and greater shelter (Orrock et al. 2004). To understand the variation in the numbers trapped of each species, we need to establish an understanding of the differing ecological requirements for each species. The two most abundant species were the wood mouse and the bank vole. The results show that wood mice are found in greater numbers in hedgerows containing standard/veteran trees. This conclusion is supported by previous studies which have shown that trees within hedgerows are beneficial for wood mice (Montgomerie and Dowie, 1993). Mice often take shelter in burrows formed beneath trees/within tree roots which may suggest why this species was found more commonly within hedgerows containing standard/veteran trees (Montgomerie and Dowie, 1993). Wood mice are a generalist species occupying a wide variety of habitat (Flowerdew 1993). They general occupy a relatively large home range and travel extensively, consuming a wide range of food sources depending upon season and availability (Flowerdew 1993). This is reflected in the results, with wood mice having been trapped in 93% of all the hedgerows. The results also show that wood mice abundance is not affected by ES management for hedgerows, nor is it significantly improved by the presence of a buffer strip. Wood mice have been shown to avoid hedgerows with major gaps, and wood mouse captures have been shown to increase with proximity to woodland (Gelling et al. 2007). Wood mice have relatively large home ranges and the suggestion is that individuals rarely stay long within any one hedgerow; rather they travel through, utilising hedgerows for foraging and shelter between woodland (Montgomery and Dowie 1993; Gelling et al 2007; Todd et al 2000; Tew et al. 2000). Therefore, ES management and the presence of buffer strips have little effect on the number of wood mouse captures; more important is the proximity to woodland or the presence of trees within a hedgerow which provide the preferred shelter for the wood mouse (Todd et al. 2000; Tew et al. 2000). Bank voles are a more specialist species, and generally occupy much smaller home ranges than do wood mice. They are burrowers, using ground vegetation to create runs and pathways in deciduous habitats (Morris 1982; Alibhai and Gipps 1985). Bank voles are a major prey resource for a number of raptors and bank vole abundance has been shown to significantly affect raptor populations (Korpimaki and Norrdahl, 1991). Other studies have found that bank vole numbers are positively associated with the size of hedgerows (Pollard Relton, 1970; Tew, 1994; Bellamy et al., 2000). Grassy margins of 2m plus have been shown to significantly increase bank vole numbers in arable fields (Shore et al. 2005), my results show that this conclusion extends to pastoral landscapes with bank vole numbers being significantly increased by the presence of an unimproved grassy margin or conservation buffer strip. The results also suggest that ES management improves hedgerows for bank voles, with bank vole abundance found to be significantly higher on ES hedgerow sites. Bank voles are found in much greater abundance in areas which provide thick ground vegetation and suffer little disturbance (Tew 1994), my results suggest that this is partially provided by ES management, however, the creation of grassy margins along hedgerows could significantly improve bank vole abundance in pastoral landscapes. The creation of margins could also be significant in the conservation of field voles. Field vole numbers in the UK are in decline believed to be due to the loss of rough grass habitat in intensively managed arable regions (Harris et al., 1995; Love et al., 2000). Field voles are specialists and depend upon rough, ungrazed grassland within woodland and hedgerows. Field voles are g enerally only found within areas of long grass (Alibhai and Gipps, 1991b). Very few captures of field voles were recorded within this experiment, however all field voles captures occurred within hedgerows flanked by conservation buffer strips. The presence of a buffer strip may provide the field voles’ preferred habitat of rough, ungrazed grassland vegetation (Alibhai and Gipps 1991b). The result is concordant with others who have found that field vole numbers can be positively associated with the presence of a conservation buffer strip (Gelling et al. 2007). Within this experiment, the size of the grassy margin is not specified beyond the requirement that it is over 2m wide from the edge of the hedgerow. Other studies have shown that field vole numbers are positively correlated with grass margin width. Margins of 6m are rarely sufficient to support viable populations of field voles (Shore et al. 2005) and individuals are rarely found in margins of 4m (Bellamy et al. 2000). F ield voles favour areas of rough grassland, specifically areas with a high proportion of long grasses and dense litter cover (Tattersall et al. 2000). With all available land being used for grazing and silaging, very few pastoral farms currently offer such wide margins with dense litter cover. With field vole numbers in the decline, there is significant scope for the improvement of pastoral farming for the benefit of field voles. Although not targeted for trapping, there were a relatively large number of common shrews caught during the two seasons of trapping. Common shrews utilise burrow and create home ranges of no more than a few hundred metres in which they spend their entire lives. They are mainly insectivorous but also eat earthworms, slugs and snails (Gurnell and Flowerdew, 1996). Common shrews were more commonly caught within hedgerows not flanked by conservation buffer strips. Common shrews are known to prefer vegetation cover for foraging (Gurnell and Flowerdew, 1996), however, my result suggests that the thick vegetation cover within buffer strips is detrimental to common shrew numbers. In the long term, the benefit of ES management for small mammals is still uncertain. Bates and Harris (2007), suggest that ES will have little benefit for overall small mammal abundance. Shore et al. (2005) show that the grassy margins included in the ES management of hedgerows in arable landscapes can have a beneficial effect on the abundance of some species. It is important to understand what factors of hedgerow management effect small mammal abundance, as this will directly influence the densities of predator species (Love et al. 2000). My results suggest that ES management does increase small mammal abundances, however, I am unable to define how ES management benefits small mammals, with the hedgerow survey drawing no definitive conclusions on how ES management is affecting hedgerow characteristics. The results of this investigation do demonstrate the value of unimproved grassy margins within pastoral landscapes. Gelling et al. (2007) also note the benefit that ‘conservation buffer strips’ have for small mammal abundances within hedgerows in pastoral landscapes. Conclusion With a large part of the British countryside being managed as agricultural land, it is important that a balance is struck between food production and creating a stable habitat that can support a diversity of wildlife. The Environmental Stewardship schemes introduced in 2005 were important as a large scale step towards environmentally sensitive farming on a national scale. However, with so much land going into ES management it is important that scientists and the government instruct farmers properly. The prescriptions laid out within the ES guidelines are now being applied to huge areas of the countryside, however, the efficacy of the schemes for the wide scale health of British wildlife remains uncertain. Correct management of farmland and therefore the information within ES guidelines is key to halting the declines seen in many species over the last few decades. There needs to be constant monitoring of the effects of ES schemes in order to know how to move forward. It is also impo rtant that scientists continue to influence the guidelines for farmers to ensure maximum benefit for British wildlife. References Alibhai, S.K., Gipps, J.H.W. (1991a) Bank vole. In: Harris, S. (ed) The handbook of British mammals. Blackwell Scientific Publishing, Oxford Alibhai, S.K., Gipps, J.H.W. (1991b) Field vole. In: Harris, S. (ed) The handbook of British mammals. Blackwell Scientific Publishing, Oxford Barr, C.J., Gillespie, M.K. (2000) Estimating hedgerow length and pattern characteristics in Great Britain using Countryside Survey data. Journal of Environmental Management, 60, 23–32 Bates, F.S., Harris, S. (2009) Does hedgerow management on organic farms benefit small mammal populations? Agriculture, Ecosystems and the Environment, 129, 124-130. Bellamy, P.E., Shore, R.F., Ardeshir, D., Treweek, J.R., Sparks, T.H. (2000) Road verges as habitat for small mammals in Britain. Mammal Review, 30, 131–139 Butet, A., Paillat, G., Delettre, Y. (2006) Seasonal changes in small mammal assemblages from field boundaries in an agricultural landscape of western France. Agriculture, Ecosystems and the Environment, 113, 364-369. DEFRA (2005a) Single Payment Scheme Cross Compliance Handbook for England 2005 Edition. DEFRA, London. Available at: https://www.defra.gov.uk/farm/capreform/pubs/pdf/Cross-compliance-3011.pdf DEFRA (2005b) Entry Level Stewardship Handbook. DEFRA, London. Available at: https://www.defra.gov.uk/erdp/pdfs/es/els-handbook.pdf DEFRA (2007) Hedgrow Survey Handbook (2nd edition). Available at: https://www.defra.gov.uk/farm/environment/landscape/documents/hedgerow-survey-handbook.pdf Fitzgibbon, C.D. (1997) Small mammals in farm woodlands: the effects of habitat, isolation and surrounding landuse patterns. Journal of Applied Ecology, 34, 530–535 Flowerdew JR (1993) Mice and voles. Whittet Books, London Flowerdew JR (1991) Wood mouse. In: Harris, S. (ed) The handbook of British mammals. Blackwell Scientific Publications, Oxford Gelling , M., Macdonald, D.W., Mathews, F. (2007) Are hedgerows the route to increased farmland small mammal density? Us e of hedgerows in British pastoral habitats. Landscape Ecology, 22, 1019–1032. Gibson, R.H., Pearce, S., Morris, R.J., Symondson, W.O.C., Memmott, J. (2007) Plant diversity and land use under organic and conventional agriculture: a whole-farm approach. Journal of Applied Ecology, 44, 792–803 Gurnell, J. Flowerdew, J.R. (1996) Live Trapping Small Mammals A Practical Guide (4th edn.) The Mammal Society, London Harris, S., Morris, P., Wray, S. Yalden, D.W. (1995) A Review of British Mammals: Population Estimates and Conservation Status of British Mammals other than Cetaceans. JNCC, Peterborough. Hayward, G.F., Phillipson, J. (1979) Community structure and functional role of small mammals in ecosystems. Ecology of Small Mammals, Chapman and Hall, London, 135–211. Hinsley, S.A., Bellamy, P.E. (2000) The influence of hedge structure, management and landscape context on the value of hedgerows to birds: A review. Journal of Environmental Management, 60, 33-49 Kl eijn, D., Sutherland, W.J. (2003) How effective are European agri-environment schemes in conserving and promoting biodiversity? Journal of Applied Ecology, 40, 947–969 Korpimaki, E., Norrdahl, K. (1991) Numerical and functional responses of kestrels, short-eared owls and long-eared owls to vole densities. Ecology, 72, 814-826 Love, R.A., Webbon, C., Glue, D.E., Harris, S. (2000) Changes in the food of British Barn Owls (Tyto alba) between 1974 and 1997. Mammal Review, 30, 107–129 Montgomerie, W.I., Dowie, M. (1993) The distribution and population regulation of the Woodmouse Apodemus sylvaticus on field boundaries of pastoral farmland. Journal of Applied Ecology, 30, 783-791. Orrock, J.L., Danielson, B.J., Brinkerhoff, R.J. (2004) Rodent foraging is affected by indirect, but not by direct, cues of predation risk. Behavioural Ecology 15, 433–437 Pocock, M.J.O., Jennings, N. (2008) Testing biotic indicator taxa: the sensitivity of insectivorous mammals and t heir prey to the intensification of lowland agriculture. Journal of Applied Ecology, 45, 151–160 Pollard, E., Relton, J. (1970) A study of small mammals in hedges and cultivated fields. Journal of Applied Ecology, 1, 549–557 Rich, T.C.G., Clements, D.K., Lewis, J., Moore, L. (2000) A comparison of four methods used to survey hedgerows: The Cardiff Hedgerow Survey 1998. Journal of Environmental Management, 60, 91–100 Robinson, R.A., Sutherland, W.J. (2002) Post-War Changes in Arable Farming and Biodiversity in Great Britain. Journal of Applied Ecology, 39, 157-176 Shore, R.F., Meek, W.R., Sparks, T.H., Pywell, R.F., Nowakowski, M. (2005) Will Environmental Stewardship enhance small mammal abundance on intensively managed farmland? Mammal Review, 35, 277–284 Soule, M.E., Terbough, J. (1999) Continental conservation: scientific foundations of regional reserve networks. Island Press, Washington, DC Tattersall, F.H., Avundo, A.E., Manley, W.J., Hart, B. J. Macdonald, D.W. (2000) Managing set-aside for field voles (Microtus agrestis). Biological Conservation, 96, 123–128 Tattersall, F.H., MacDonald, D.W., Hart, B.J., Johnson, P., Manley, W., Feber, R. (2002) Is habitat linearity important for small communities on farmland. Journal of Applied Ecology, 39, 643-652. Tattersall, F.H., Macdonald, D.W., Hart, B.J., Manley, W.J. (2004) Balanced dispersal or source-sink – do both models describe wood mice in farmed landscapes? Oikos, 106, 536–550 Tattersall, F.H., Macdonald, D.W., Hart, B.J., Manley, W.J., Feber, R.E. (2001) Habitat use by wood mice (Apodemus sylvaticus) in a changeable arable landscape. Journal of Zoology, 255, 487–494 Tew TE, Todd IA, Macdonald DW (2000) Arable habitat use by wood mice (Apodemus sylvaticus). 2. Microhabitat. Journal of Zoology, 250, 305–311 Tew, T.E., Todd, I.A. MacDonald, D.W. (1994) Field margins and small mammals. In: Field Margins: Integrating Agriculture and Conservation, 85–94. British Crop Protection Council, Surrey. Todd, I.A., Tew, T.E., Macdonald, D.W. (2000) Arable habitat use by wood mice (Apodemus sylvaticus). 1. Macrohabitat. Journal of Zoology, 250, 299–303